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Inglorious Basterds is set in an alternate history of the Second World War in which the entire top leadership of Nazi Germany, namely Hitler, Goering, Goebbels and Bormann attend a film premiere in Paris celebrating the exploits of a German sniper who had managed to kill 250 American soldiers in Italy. The timeframe of the film is set in early June 1944, after the D-Day landings but before the liberation of Paris.

The film tracks the separate attempts to kill Hitler by two disparate forces, one being the "Basterds", a motley crew of Jewish American soldiers out for revenge against the Nazis. The Basterds have a modus operandi whereby each soldier must cut off the scalp of a dead Nazi soldier. The Basterds allow one German soldier to survive in order to spread the news of the terror of their attacks. However, the Basterds carve a swastika into the forehead of that German. The other force concerns Shosanna, the only Jewish survivor of an attack led by the "Jew Hunter" that had killed all her family. The Basterds and Shosanna remain unaware of each other throughout the film.